Wolves, bears, and man are the primary predators of muskoxen. They are also vulnerable to accidents, such as falling from cliffs or drowning, and starvation if deep snow or ice covers their food. Cows may live more than 20 years, but the average lifespan is much less. On the average, bulls probably die at a younger age than cows due to the increased risks during fights over females. Muskox populations were extirpated from Alaska in the late 1800s, and apparently declined in Canada and Greenland by the early 20th Century. Although scientists are not sure about the exact causes of the decline, hunting and climatic changes may have been factors.